"Every perception is to some degree an act of creation, and every act of memory is to some degree an act of imagination."

-- Gerald Edelman, Second Nature: Brain Science and Human Knowledge
Part of developing Anki as a virtuoso skill is cultivating the ability to use it for types of understanding beyond basic facts. Indeed, many of the observations I've made (and will make, below) about how to use Anki are really about what it means to understand something. Break things up into atomic facts. Build rich hierarchies of interconnections and integrative questions. Don't put in orphan questions. Patterns for how to engage with reading material. Patterns (and anti-patterns) for question types. Patterns for the kinds of things you'd like to memorize. Anki skills concretely instantiate your theory of how you understand; developing those skills will help you understand better. It's too strong to say that to be a virtuoso Anki user is to be a virtuoso in understanding. But there's some truth to it.